Besides the meibomian glands, which accessory glands also secrete oil?

The correct answer identifies the Zeiss and Moll glands as additional accessory glands that secrete oil.

The Zeiss glands are specialized sebaceous glands located at the base of the eyelashes, and they produce a lipid secretion that helps to lubricate the eyelashes and contribute to the overall tear film stability. The Moll glands, on the other hand, are modified sweat glands located in the eyelid margins, also contributing lipid to the tear film. Together, these glands play a significant role in maintaining the health and function of the ocular surface by preventing evaporation of the tears and providing a protective barrier.

Understanding the roles of these glands emphasizes the complexity of the tear film and the importance of lipid secretions in ocular health, which can be particularly relevant in the context of dry eye disease and other ocular surface disorders.