In addition to the meibomian glands, which other accessory glands secrete oil?

The meibomian glands are known for producing the lipid layer of the tear film, essential for reducing evaporation of tears. In addition to these glands, the Zeiss and Moll glands also contribute to the secretion of oil. Zeiss glands are located at the base of the eyelashes and secrete an oily substance that helps to lubricate the eyelash follicles, thereby assisting in the overall maintenance of the eyelid and eye surface. Similarly, Moll glands, which are modified sweat glands found in the eyelids, also produce a type of oily secretion that contributes to the tear film and enhances lubrication.

This complementary oil secretion from both the Zeiss and Moll glands plays a crucial role in maintaining the health of the ocular surface by providing protection and preventing tear film instability. The other combinations of accessory glands listed do not represent glands that primarily secrete oil in conjunction with the meibomian glands.